Powdermill Avian Research Center
Fall 2004
Bird Banding Summary




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Powdermill Nature Reserve's Bird Banding Program, located on the 2,200-acre biological field station of Carnegie Museum of Natural History in the Laurel Highlands region of southwestern Pennsylvania, completed its 43rd fall migration banding season in 2004.  The station was in operation 99 out of a possible 120 days from August through November.  Total banding effort for the fall 2004 season was 27,670 net-hours (average 231 net/trap hrs./day), the first of many record setting highs for this fall.  Far exceeding the 42-year average of 15,373 net hours, this total also exceeded our previous banding effort record of 23,655 by more than 4,000 hours and surely contributed to our above average banding total of 7798 birds (42 year avg. 6298). 
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The large number of net hours this fall reflects both favorable weather conditions for banding throughout much of the season and the addition of a few more nets around the banding area, but, it is attributable to the assistance and dedication of all of our visting bander friends and colleagues, interns, volunteers, and fall bander workshop participants. 

For this reason we begin by saying THANK YOU to: Mike Comley, Bill Diehl, Pam Ferkett, Randi Gerrish, Tom Greg, Dan Hinnebusch, Jim Ingold, Brian Jones, Carroll Labarthe, David Liebman, Annie Lindsay, Cokie Lindsay, Gary and Sandy Lockerman, Molly McDermott, Trish Miller, Jake Mohlmann, Felicity Newell, Robert Nugent, Kelly Perkins, Katherine Reinamen, Joe Schreiber, Carole Shanahan, Debbie and Al Sherkow, Matt Shumar, Rosemary Spreha, and Guy Ubaghs. (names in red are those who volunteered a week or more of their time to the fall banding efforts at Powdermill this year)

We benefited once again from an extended visit by Dr. David Norman, our British friend and ringing colleague from the Merseyside Ringing Group, whose visit this fall, again, coincided perfectly for adding assistance and some "overseas" banding insight during part of the first fall bander workshop offered this year.  We were also fortunate enough to be paid a visit by the Bird Banding Laboratory's recent acting chief and long-time biologist, Mary Gustafson.

We also thank our workshop participants Carol and Fred McCullogh, Deb Plotts, Gretchen Putonen, Clay Corbin, Fred Bassett, Virginia Duffy, and Tom LeBlanc for their participation and especially their help and patience during the "busy days" when normal workshop activities were put on hold. 

Finally,we are extremely fortunate to have Dr. David Smith as the director of Powdermill Nature Reserve and in addition to the help he's provided on more than one occassion to the general daily banding operations, we are  grateful for his unfailing support of the banding program and research at the Powdermill Avian Research Center.  Last, but certainly not least, we thank our highly skilled data entry operator at Carnegie Museum's Section of Birds, Marilyn Niedermeier, who painstakingly enters, checks and double checks all of our banding records, and who is responsible for generating banding schedules and filling data requests. 

Summary

Our total of 7,798 is, not counting our record fall total in 2001, the highest fall banding total since 1995.  We banded 100 or more birds on 30 days during the season (compared to 24 days last fall).  Peak banding dates were September 29 (307), September 14 (291) and October 24 (240).  Eighteen percent of our cumulative total was banded during August, 40% in September, 32% in October, and 10% in November (top graph below).  Note also that our highest capture rates were on days with the lowest banding totals and reflect effort from only a few localized nets targeting birds at feeding stations.

Importantly, the record high banding effort this fall was not matched by a comparatively very large (albeit above average) banding total, such that the calculated capture rate this season (28.2 birds/100 net-hours) was far below average.  Clearly, in examining possible long-term trends in fall banding data, it is necessary to account for variation in banding effort across years (bottom graph below).
 

ChartObject Chart 1

ChartObject Powdermill Avian Research Center Fall Net Hours vs. Capture RatesCorrelation r = -.62 (P < .001)


The closely corresponding relationship between effort and totals is confirmed when we examine the correlation between these two variables in the graph below.  Note that the banding total for fall 2004 (green symbol) is slightly lower than would be predicted from the overall correlation given the higher than average banding effort this fall.

ChartObject Powdermill Avian Research Center  Fall Banding Effort vs. Banding TotalsCorrelation r =.52 (P < .001)

This fall 110 species of birds were banded, which ties as the fifth highest ever species total for the fall season.   (click here for a table of species totals given in descending order).  The diversity of species banded by day is plotted in the graph below.  Unlike most years at Powdermilll where species diversity peaks two to three weeks before banding totals, the days with the most species banded (47 and 43), also coincided with the highest daily banding totals for the season (on 9/14 and 9/29 respectively). 

ChartObject Chart 4


Our Fall 2004 season highlight, undoubtedly, was the banding of the first ever Black-headed Grosbeak in the 43 year history of the program, bringing the cumulative Powdermill fall banding list to 172 species (including two hybrid forms and two recognizable subspecies). 

A few other good "gets" (species for which we average less than one banded per fall) included: Green Heron, Spotted Sandpiper, Pileated Woodpecker, Northern Mockingbird, Worm-eating Warbler, and Grasshopper Sparrow.

Altenatively, conspicuous "misses" this fall (i.e., species for which an average of one or more has been banded during fall) included: Red-winged Blackbird, Barn Swallow, Mourning Dove, Evening Grosbeak, Olive-sided Flycatcher, Prairie Warbler, House Sparrow, and Marsh Wren.

Highs and Lows


Eight species set record high totals this fall, many surpassing the previous high totals by 50 or more birds.  Black-throated Green Warbler crushed the previous record of 130 with 208 birds banded and Hooded Warbler followed in suit with 237 beating out the previous record of 189.  Red-eyed Vireo surpassed the record total set just last year of 249 with 303 birds.  The remaining species records are as follows with 2004 fall total/previous record high: Acadian Flycatcher 44/42, Chestnut-sided Warbler 101/98, Black-throated Blue Warbler 101/74, Scarlet Tanager 145/108, and Blue-headed Vireo 74/53.  These record totals, many of locally breeding bird species, seem to be further evidence of the very productive 2004 breeding season experienced by many birds in the Laurel Valley, as well as elsewhere.  See also the 2004 Summer Summary

In the tables below, totals and capture rates for fall 2004 are compared to averages, standard deviations (S.D.), minima, and maxima for the preceding 42 years.  White rows highlight species that were one or more S.D. above or below the long-term average for banding total and capture rate (comparisons were made only for species with a long-term average banding total >1).  Species in red were below average; blue denotes species that were above average. 

An incredible 29 species were one or more standard deviations above average in fall 2004, in terms of raw banding total, and only two were below average.  Confirming the importance of adjusting for effort in comparing banding totals from year to year, examination of capture rates gives a very different picture with just eight species being above average and 15 below average. 


 
 

Species Total Number of Birds Banded
  Fall Seasons, 1962-2003
2004 AVE SD MIN MAX
Green Heron 1 0.7 1.0 0 4
Sharp-shinned Hawk 3 2.1 2.2 0 9
Solitary Sandpiper 4 2.6 3.3 0 14
Spotted Sandpiper 1 0.3 0.7 0 4
American Woodcock 2 1.6 1.5 0 6
Mourning Dove 0 2.3 4.1 0 23
Black-billed Cuckoo 3 3.0 3.2 0 12
Yellow-billed Cuckoo 1 1.7 1.9 0 8
Eastern Screech-Owl 1 0.6 0.8 0 4
Northern Saw-whet Owl 19 0.5 1.0 0 4
Ruby-throated Hummingbird 299 125.1 68.2 30 309
Belted Kingfisher 1 4.1 2.8 0 10
Red-bellied Woodpecker 3 1.0 1.9 0 9
Yellow-bellied Sapsucker 7 3.6 2.6 0 11
Downy Woodpecker 15 14.9 6.2 5 34
Hairy Woodpecker 1 1.7 1.1 0 5
Northern Flicker 5 7.3 4.6 0 19
Pileated Woodpecker 1 0.0 0.3 0 2
Olive-sided Flycatcher 0 1.9 2.4 0 11
Eastern Wood-Pewee 40 29.8 16.2 5 66
Yellow-bellied Flycatcher 81 60.4 26.5 17 121
Acadian Flycatcher 44 13.2 8.7 0 42
Traill's (Willow and Alder) Flycatcher 34 20.8 8.9 5 39
Least Flycatcher 64 89.0 43.6 32 191
Eastern Phoebe 135 60.5 34.9 19 202
Great-crested Flycatcher 1 1.7 1.5 0 6
Eastern Kingbird 0 0.9 1.5 0 7
White-eyed Vireo 4 5.9 5.1 0 16
Yellow-throated Vireo 7 1.8 1.7 0 6
Blue-headed Vireo 74 21.1 13.2 2 53
Warbling Vireo 1 1.1 0.9 0 3
Philadelphia Vireo 21 20.4 14.1 2 69
Red-eyed Vireo 303 126.4 48.5 46 249
Blue Jay 24 17.0 12.8 0 68
Barn Swallow 0 4.5 5.3 0 20
Black-capped Chickadee 177 128.5 115.5 23 490
Tufted Titmouse 18 27.0 11.5 8 52
Red-breasted Nuthatch 0 0.4 0.7 0 3
Black-headed Grosbeak 1        
White-breasted Nuthatch 7 7.1 3.5 2 19
Brown Creeper 8 7.1 3.7 2 16
Carolina Wren 8 4.6 6.0 0 26
House Wren 23 50.5 27.4 8 122
Winter Wren 36 20.4 16.2 1 66
Marsh Wren  0 2.0 2.4 0 13
Golden-crowned Kinglet 65 42.8 26.6 4 119
Ruby-crowned Kinglet 298 205.7 97.5 37 441
Blue-gray Gnatcatcher 7 6.7 4.1 0 16
Eastern Bluebird 1 3.5 5.0 0 18
Veery 14 10.5 6.5 1 25
Gray-cheeked Thrush 46 40.6 33.3 5 153
Swainson's Thrush 274 215.7 149.0 36 619
Hermit Thrush 46 47.8 22.3 11 116
Wood Thrush 119 28.2 22.7 3 120
American Robin 20 19.6 15.6 0 63
Gray Catbird 284 240.0 73.8 95 390
Northern Mockingbird 1 0.2 0.5 0 2
Brown Thrasher 4 8.6 6.8 0 32
European Starling 0 0.5 1.2 0 5
Cedar Waxwing 174 262.5 266.9 56 1235
Blue-winged Warbler 20 6.5 5.7 0 23
Golden-winged Warbler 1 3.0 2.3 0 10
"Brewster's" Warbler 0 0.4 0.7 0 2
Tennessee Warbler 60 140.7 149.8 8 729
Orange-crowned Warbler 2 5.9 5.2 0 25
Nashville Warbler 28 48.1 21.5 13 101
Northern Parula 9 2.1 2.0 0 10
Yellow Warbler 3 5.8 3.7 1 18
Chestnut-sided Warbler 101 40.0 22.2 6 98
Magnolia Warbler 322 225.5 96.6 62 523
Cape May Warbler 28 88.3 88.1 8 365
Black-throated Blue Warbler 101 21.9 17.3 1 74
Yellow-rumped Warbler 304 307.2 206.2 58 979
Black-throated Green Warbler 208 53.4 29.6 15 130
Blackburnian Warbler 8 7.9 6.1 0 25
Pine Warbler 0 0.2 0.5 0 2
Prairie Warbler 0 2.6 2.5 0 9
"Western" Palm Warbler 21 32.8 26.9 4 145
Bay-breasted Warbler 12 28.3 30.1 1 113
Blackpoll Warbler 33 32.4 15.2 11 92
Cerulean Warbler 1 1.5 1.6 0 6
Black-and-white Warbler 13 14.4 7.6 1 32
American Redstart 165 110.8 52.4 19 222
Worm-eating Warbler 1 0.8 1.1 0 5
Ovenbird 64 54.3 24.4 18 120
Northern Waterthrush 52 31.7 14.9 6 71
Louisiana Waterthrush 3 1.8 1.7 0 6
Kentucky Warbler 8 8.5 4.4 1 22
Connecticut Warbler 10 16.1 7.1 5 31
Mourning Warbler 11 13.6 6.1 5 27
Common Yellowthroat 226 245.0 68.2 90 427
Hooded Warbler 237 71.4 44.6 14 189
Wilson's Warbler 20 30.6 15.0 14 86
Canada Warbler 43 28.9 14.1 6 74
Yellow-breasted Chat 0 4.2 4.5 0 19
Scarlet Tanager 145 47.9 25.3 2 108
Eastern Towhee 59 42.7 19.2 13 85
American Tree Sparrow 14 26.1 19.9 4 99
Chipping Sparrow 30 45.9 40.7 6 172
Field Sparrow 54 141.5 97.5 33 394
Savannah Sparrow 3 2.0 2.7 0 14
Grasshopper Sparrow 1 0.6 1.4 0 7
Henslow's Sparrow 0 0.2 0.4 0 1
Fox Sparrow 78 34.0 20.9 7 95
Song Sparrow 322 304.5 84.4 171 511
Lincoln's Sparrow 46 59.1 23.7 22 130
Swamp Sparrow 161 156.7 54.6 55 294
White-throated Sparrow 450 366.6 177.3 110 875
White-crowned Sparrow 14 15.9 8.9 3 50
Dark-eyed Junco 359 409.3 191.5 86 945
Northern Cardinal 53 39.3 11.5 22 68
Rose-breasted Grosbeak 156 64.6 36.0 2 140
Indigo Bunting 99 68.9 29.1 16 134
Red-winged Blackbird 0 6.4 9.4 0 48
Rusty Blackbird 6 14.9 18.4 0 94
Common Grackle 1 3.4 9.7 0 50
Brown-headed Cowbird 1 1.0 1.6 0 7
Baltimore Oriole 2 2.7 2.6 0 9
Purple Finch 92 173.8 219.1 0 911
House Finch 7 55.6 83.5 0 395
Pine Siskin 2 32.9 64.8 0 361
American Goldfinch 693 397.7 243.0 160 1654
Evening Grosbeak 0 31.5 87.7 0 511
House Sparrow 0 31.8 36.7 0 143
           
Species Capture Rates (no. birds banded/1000 net hrs.)
  Fall Seasons, 1962-2003
2004 AVE SD MIN MAX
Green Heron 0.0 0.0 0.1 0.0 0.3
Sharp-shinned Hawk 0.1 0.1 0.1 0.0 0.5
Solitary Sandpiper 0.1 0.2 0.2 0.0 0.7
Spotted Sandpiper 0.0 0.0 0.1 0.0 0.6
American Woodcock 0.1 0.1 0.1 0.0 0.4
Mourning Dove 0.0 0.1 0.2 0.0 1.3
Black-billed Cuckoo 0.1 0.2 0.3 0.0 1.3
Yellow-billed Cuckoo 0.0 0.1 0.2 0.0 0.8
Eastern Screech-Owl 0.0 0.0 0.1 0.0 0.2
Northern Saw-whet Owl 0.7 0.0 0.1 0.0 0.2
Ruby-throated Hummingbird 10.8 8.4 4.6 2.2 22.2
Belted Kingfisher 0.0 0.3 0.2 0.0 0.6
Red-bellied Woodpecker 0.1 0.1 0.1 0.0 0.6
Yellow-bellied Sapsucker 0.3 0.2 0.2 0.0 0.6
Downy Woodpecker 0.5 1.0 0.4 0.4 2.3
Hairy Woodpecker 0.0 0.1 0.1 0.0 0.4
Northern Flicker 0.2 0.5 0.3 0.0 1.5
Pileated Woodpecker 0.0 0.0 0.0 0.0 0.1
Olive-sided Flycatcher 0.0 0.1 0.2 0.0 0.8
Eastern Wood-Pewee 1.4 2.0 1.0 0.4 4.4
Yellow-bellied Flycatcher 2.9 4.0 1.7 1.4 6.9
Acadian Flycatcher 1.6 0.9 0.7 0.0 4.2
Traill's (Willow and Alder) Flycatcher 1.2 1.4 0.5 0.5 2.4
Least Flycatcher 2.3 6.2 3.7 2.2 18.6
Eastern Phoebe 4.9 4.0 2.1 1.4 11.1
Great-crested Flycatcher 0.0 0.1 0.1 0.0 0.6
Eastern Kingbird 0.0 0.1 0.2 0.0 1.1
White-eyed Vireo 0.1 0.4 0.4 0.0 1.5
Yellow-throated Vireo 0.3 0.1 0.1 0.0 0.4
Blue-headed Vireo 2.7 1.4 0.9 0.1 4.3
Warbling Vireo 0.0 0.1 0.1 0.0 0.2
Philadelphia Vireo 0.8 1.4 1.0 0.2 4.3
Red-eyed Vireo 11.0 8.9 4.7 3.1 22.4
Blue Jay 0.9 1.1 0.8 0.0 3.7
Barn Swallow 0.0 0.3 0.4 0.0 1.6
Black-capped Chickadee 6.4 8.8 8.1 1.2 34.8
Tufted Titmouse 0.7 1.8 0.7 0.6 3.5
Red-breasted Nuthatch 0.0 0.0 0.0 0.0 0.2
Black-headed Grosbeak 0.0        
White-breasted Nuthatch 0.3 0.5 0.2 0.1 1.1
Brown Creeper 0.3 0.5 0.2 0.1 1.0
Carolina Wren 0.3 0.3 0.4 0.0 1.6
House Wren 0.8 3.3 1.5 1.1 7.5
Winter Wren 1.3 1.3 0.9 0.2 4.1
Marsh Wren  0.0 0.1 0.1 0.0 0.7
Golden-crowned Kinglet 2.3 2.7 1.6 0.4 6.8
Ruby-crowned Kinglet 10.8 13.5 6.9 5.5 45.9
Blue-gray Gnatcatcher 0.3 0.4 0.3 0.0 1.0
Eastern Bluebird 0.0 0.3 0.4 0.0 1.6
Veery 0.5 0.7 0.4 0.1 2.1
Gray-cheeked Thrush 1.7 2.8 2.3 0.4 11.8
Swainson's Thrush 9.9 14.4 9.6 3.9 36.5
Hermit Thrush 1.7 3.2 1.6 1.4 9.0
Wood Thrush 4.3 1.8 1.2 0.2 5.6
American Robin 0.7 1.3 1.1 0.0 4.3
Gray Catbird 10.3 16.5 6.5 5.9 36.5
Northern Mockingbird 0.0 0.0 0.0 0.0 0.1
Brown Thrasher 0.1 0.6 0.5 0.0 2.6
European Starling 0.0 0.0 0.1 0.0 0.3
Cedar Waxwing 6.3 19.5 27.9 3.8 172.6
Blue-winged Warbler 0.7 0.4 0.4 0.0 1.6
Golden-winged Warbler 0.0 0.2 0.2 0.0 0.9
"Brewster's" Warbler 0.0 0.0 0.0 0.0 0.1
Tennessee Warbler 2.2 9.4 9.1 0.6 37.0
Orange-crowned Warbler 0.1 0.4 0.3 0.0 1.1
Nashville Warbler 1.0 3.2 1.3 1.2 6.9
Northern Parula 0.3 0.1 0.1 0.0 0.6
Yellow Warbler 0.1 0.4 0.3 0.1 1.8
Chestnut-sided Warbler 3.7 2.7 1.4 0.5 6.2
Magnolia Warbler 11.6 14.7 5.0 6.0 24.4
Cape May Warbler 1.0 6.1 6.2 0.4 26.4
Black-throated Blue Warbler 3.7 1.4 1.0 0.1 4.1
Yellow-rumped Warbler 11.0 20.2 12.4 4.2 56.7
Black-throated Green Warbler 7.5 3.5 1.6 1.1 7.0
Blackburnian Warbler 0.3 0.5 0.4 0.0 2.0
Pine Warbler 0.0 0.0 0.0 0.0 0.1
Prairie Warbler 0.0 0.2 0.2 0.0 0.8
"Western" Palm Warbler 0.8 2.2 1.9 0.3 9.2
Bay-breasted Warbler 0.4 1.8 1.8 0.1 6.8
Blackpoll Warbler 1.2 2.2 0.9 0.8 5.2
Cerulean Warbler 0.0 0.1 0.1 0.0 0.4
Black-and-white Warbler 0.5 1.0 0.7 0.1 4.0
American Redstart 6.0 7.3 3.3 1.5 14.9
Worm-eating Warbler 0.0 0.1 0.1 0.0 0.4
Ovenbird 2.3 3.6 1.5 1.2 7.3
Northern Waterthrush 1.9 2.1 1.0 0.6 5.2
Louisiana Waterthrush 0.1 0.1 0.1 0.0 0.5
Kentucky Warbler 0.3 0.6 0.4 0.1 1.8
Connecticut Warbler 0.4 1.1 0.5 0.4 2.5
Mourning Warbler 0.4 0.9 0.3 0.3 1.7
Common Yellowthroat 8.2 16.3 4.3 9.1 25.8
Hooded Warbler 8.6 4.7 2.6 0.8 9.6
Wilson's Warbler 0.7 2.0 0.8 0.9 4.2
Canada Warbler 1.6 1.9 1.0 0.5 5.2
Yellow-breasted Chat 0.0 0.3 0.4 0.0 2.6
Scarlet Tanager 5.2 3.5 2.5 0.1 13.0
Eastern Towhee 2.1 2.9 1.5 1.1 9.6
American Tree Sparrow 0.5 1.8 1.5 0.1 5.9
Chipping Sparrow 1.1 3.0 2.7 0.5 12.6
Field Sparrow 2.0 9.4 6.0 2.1 25.5
Savannah Sparrow 0.1 0.2 0.3 0.0 2.1
Grasshopper Sparrow 0.0 0.1 0.2 0.0 1.1
Henslow's Sparrow 0.0 0.0 0.0 0.0 0.2
Fox Sparrow 2.8 2.3 1.5 0.6 7.6
Song Sparrow 11.6 20.8 7.6 9.7 50.2
Lincoln's Sparrow 1.7 4.0 2.0 1.7 12.6
Swamp Sparrow 5.8 10.3 3.0 5.2 17.8
White-throated Sparrow 16.3 24.5 11.3 7.5 53.8
White-crowned Sparrow 0.5 1.1 0.6 0.3 3.6
Dark-eyed Junco 13.0 27.6 13.0 8.2 56.4
Northern Cardinal 1.9 2.6 0.8 1.3 4.7
Rose-breasted Grosbeak 5.6 4.3 2.4 0.1 8.9
Indigo Bunting 3.6 4.8 2.7 0.9 13.1
Red-winged Blackbird 0.0 0.4 0.5 0.0 2.3
Rusty Blackbird 0.2 1.1 1.7 0.0 9.8
Common Grackle 0.0 0.2 0.6 0.0 2.8
Brown-headed Cowbird 0.0 0.1 0.1 0.0 0.6
Baltimore Oriole 0.1 0.2 0.3 0.0 1.4
Purple Finch 3.3 11.2 13.2 0.0 43.6
House Finch 0.3 4.1 6.6 0.0 28.2
Pine Siskin 0.1 2.3 4.4 0.0 24.8
American Goldfinch 25.0 26.6 13.9 9.9 77.2
Evening Grosbeak 0.0 2.5 7.4 0.0 41.1
House Sparrow 0.0 2.1 2.5 0.0 11.0